Williams TN, Mwangi TW, Wambua S, Peto TE, Weatherall DJ, Gupta S, Recker M, Penman B, Uyoga S, Mwacharia A, Mwacharo JK, Snow RW, Marsh K(2005). Negative epistasis between the malaria protective effects of alpha+-thalassemia and the sickle cell trait. Nature Genetics, 37: 1253-1257[PMCID: 3521056] ,[PMID: 16227994]